Bygone Russian royal

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TSAR.

Last seen on: Daily Boston Globe Crossword Answers Friday, 21 April 2023

Random information on the term “TSAR”:

The Stramenopiles, also called Heterokonts, are a clade of organisms distinguished by the presence of stiff tripartite external hairs. In most species, the hairs are attached to flagella, in some they are attached to other areas of the cellular surface, and in some they have been secondarily lost (in which case relatedness to stramenopile ancestors is evident from other shared cytological features or from genetic similarity). Stramenopiles represent one of the three major clades in the SAR supergroup, along with Alveolata and Rhizaria.

Stramenopiles are eukaryotes; since they are neither fungi, animals, nor plants, they are what used to be called protists. Most stramenopiles are single-celled, but some are multicellular algae including some large seaweeds, the brown algae. The group includes a variety of algal protists, heterotrophic flagellates, opalines and closely related proteromonad flagellates (all endobionts in other organisms); the actinophryid heliozoa, and oomycetes. The tripartite hairs have been lost in some stramenopiles – for example in most diatoms (although these organisms still express mastigonemic proteins – see below).

TSAR on Wikipedia